Mollusca and malacology in Canada
2024
Forsyth RG ("2023") [2024] Terrestrial molluscs of Nepisiguit Protected Natural Area and Mount Carleton Provincial Park, north-central New Brunswick. The Canadian Field-Naturalist 137 (1–2): 87–95. doi: 10.22621/cfn.v137i1.3051 · Link to PDF
No comprehensive study of the terrestrial gastropod fauna of northern New Brunswick has previously been undertaken. Most terrestrial gastropod molluscs are tiny and generally live unnoticed under debris, in leaf litter, and among other dead and living vegetation. Sifting and sorting collected samples of leaf litter are effective in sampling snails. In June–July 2015 and August 2016, 43 samples of forest leaf litter were collected in Nepisiguit Protected Natural Area and Mount Carleton Provincial Park. These samples yielded 22 species of terrestrial gastropods. Seven species are reported from New Brunswick for the first time.
Aucune étude approfondie de la faune de gastéropodes terrestres du nord du Nouveau-Brunswick n’a été entreprise auparavant. La plupart des mollusques gastéropodes terrestres sont minuscules et vivent généralement inaperçus sous les débris, dans la litière de feuilles et d’autres végétaux morts et vivants. Le tamisage et le tri des échantillons prélevés de litière de feuilles sont efficaces pour échantillonner les escargots. En juin–juillet 2015 et en août 2016, 43 échantillons de litière de feuilles forestières ont été prélevés dans l’zone naturelle protégée Nepisiguit et le parc provincial Mont-Carleton. Ces échantillons ont donné 22 espèces de gastéropodes terrestres. Sept espèces sont rapportées au Nouveau-Brunswick pour la première fois.
2022
Forsyth RG, Arsenault MA, Curley R (2022) Leaf-litter-dwelling microsnails of Prince Edward Island, Canada (Mollusca, Gastropoda, Eupulmonata). Check List 18 (5): 1005–1015. doi: 10.15560/18.5.1005
Samples of leaf litter were collected from 22 woodland sites throughout Prince Edward Island, Canada. Eighty-two specimens belonging to 15 species of litter-dwelling land snails were recovered from 20 of the 22 samples. One-third of the species—Carychium exile H.C. Lea, 1842, Helicodiscus shimeki Hubricht, 1982, Strobilops labyrinthicus (Say, 1817), Perpolita binneyana (Morse, 1864), and Striatura ferrea Morse, 1864—were previously undocumented in the literature as occurring in PEI. Leaf-litter sampling, even in smaller quantities, is an effective method for finding the often-neglected terrestrial mollusc fauna, which is composed largely of minute species.
2021
COSEWIC (2021) COSEWIC assessment and status report on the Black Hills Mountainsnail Oreohelix cooperi in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa, Ontario, Canada. Link to PDF [Status report written by R.G. Forsyth]
2020
Silva FS, Forsyth RG, Salvador RB (2020) Helicodiscus theresa from Brazil is the exotic species Helicodiscus parallelus (Gastropoda, Helicodiscidae). Strombus 26 (1–2): 15–18. Link to publication
Nicolai A, Forsyth RG (2020) Introduced Prophysaon andersonii (J.G. Cooper, 1872) in Quebec, Canada: first record of Prophysaon (Gastropoda, Eupulmonata, Arionoidea) in eastern North America, confirmed by partial-COI gene sequence. Check List 16 (2): 307–316. doi: 10.15560/16.2.307
We report for the first time the terrestrial slug Prophysaon andersonii (J.G. Cooper, 1872) from Quebec, Canada. Two specimens were collected in Parc national du Bic. The identification was determined by the external morphology and partial-COI gene sequence data. The genus Prophysaon is endemic to western North America, and the new record indisputably represents an introduction. No species of Prophysaon has, until now, been noticed in North America from outside its native range.
Nicolai A, Forsyth RG, Grantham M, Hamel CD ("2019") [2020] Tall grass prairie ecosystem management—a gastropod perspective. The Canadian Field-Naturalist 133 (4): 313–324. doi: 10.22621/cfn.v133i4.2217
Less than 5% of the original tall grass prairie in North America remains. A portion of this remnant, composed of wetland, grassland and forest, is protected by the Nature Conservancy of Canada (NCC) in southern Manitoba. This heterogeneous ecosystem has rich biodiversity; however, gastropods have not been surveyed in Canada’s tall grass prairie. We studied gastropods in Prairie, Wet Meadow, Forest, and Wet Forest habitats of the Manitoba Tall Grass Prairie Preserve that vary with respect to land management practices (prescribed burning, grazing by cattle). Gastropod community composition was unique in the Prairie where mounds of grass litter form permanently moist cavities harbouring aquatic species, while dry-habitat species colonized the upper parts of these mounds. Gastropod communities in Prairie habitats were negatively affected by grazing and burning that occurred in the five years prior to our survey. Unburned Forest patches included both forest gastropod species and edge effect influenced open-habitat species and harboured the most diverse gastropod communities. These unburned Forest patches potentially provide a species pool for post-burn prairie recolonization. The gastropod community of Wet Meadows was not affected by grazing and was composed mainly of aquatic species. In this gastropod survey five species were recorded from Manitoba for the first time. The rare Blade Vertigo (Vertigo milium) is also reported.
2019
Forsyth RG, Kamstra J (2019) Roman Snail, Helix pomatia (Mollusca: Helicidae), in Canada. The Canadian Field-Naturalist 133 (2): 156–159. doi: 10.22621/v133i2.2150
Populations of Roman Snail, Helix pomatia, a large European land snail, are reported for the first time in Canada from disturbed habitats in two distant locations: Sarnia, Ontario and Montrose, British Columbia. As Roman Snail is an edible species subject to international commercial trade, its deliberate, but illegal, introduction into Canada and intentional or unintentional releases are possible sources of these populations.
Nicolai A, Forsyth RG (2019) First record of Vallonia costata (O.F. Müller, 1774) (Gastropoda, Eupulmonata, Valloniidae) from British Columbia, Canada, confirmed by partial-COI gene sequence. Check List 15 (2): 287–293. doi: 10.15560/15.2.287
The minute terrestrial snail Vallonia costata (O.F. Müller, 1774) is reported for the first time from the province of British Columbia, Canada. The identification was based on shell morphology and confirmed by COI gene data. This species is presumed to be introduced to the province.
2018
Nicolai A, Forsyth RG (2018) Guppya sterkii (Dall, 1888) (Gastropoda, Eupulmonata, Euconulidae) in eastern Ontario, Canada. Check List 14 (4): 579–583. doi: 10.15560/14.4.579
A seldom-collected terrestrial snail, Guppya sterkii (Dall, 1888), is recorded for the first time from an older-growth hardwood forest in rural Ottawa, eastern Ontario. This represents a range extension of roughly 175 km north-east of the nearest previously known occurrence. Its conservation status and possible threats are briefly discussed.
2017
Forsyth RG (2017) On the anatomy of Novisuccinea strigata (L. Pfeiffer, 1855) (Gastropoda, Stylommatophora, Succineidae) from British Columbia, Canada. Basteria 81(4–6): 123–128. Link to publication
Novisuccinea strigata (L. Pfeiffer, 1855) is an arctic-boreal terrestrial mollusc that is both amphiberingian and occurring across a large area of northwestern Canada, including northern British Columbia, Yukon, and the Northwest Territories. The shell, jaw, and reproductive anatomy is described from specimens collected near the Hyland River, on the Liard Plain for northern British Columbia. Reproductive anatomy, and most notably the twisting of the free oviduct around the duct of the bursa copulatrix, indicates that this species properly belongs to the genus Novisuccinea as redefined by some American workers.
2016
Forsyth RG, Maunder JE, McAlpine DF, Noseworthy RG (2016) Distributional status of an introduced land snail Discus rotundatus (Rotund Disc, Mollusca: Discidae) in Canada. The Canadian Field-Naturalist 130 (3): 235–246. doi: 10.22621/cfn.v130i3.1887
First collected in North America in 1937 on the Avalon Peninsula of the Island of Newfoundland, the introduced, primarily European land snail, Discus rotundatus, has now been recorded from the Island of Newfoundland, Nova Scotia, New Brunswick, Quebec, Ontario, and British Columbia. we review all known records from Canada, demonstrate that D. rotundatus is more widespread than was previously recognized on the Island of Newfoundland, and report the first record from New Brunswick.
Forsyth RG, Catling P, Kostiuk B, McKay-Kuja S, Kuja A (2016) Pre-settlement snail fauna on the Sandbanks baymouth bar, Lake Ontario, compared with nearby contemporary faunas. The Canadian Field-Naturalist 130 (2): 152–157. doi: 10.22621/cfn.v130i2.1839
The terrestrial snail fauna in a pre-settlement soil layer, radiocarbon dated at approximately 1065–1560 years old, on the baymouth bar at Sandbanks Provincial Park, Lake Ontario, was compared with 4 nearby contemporary snail faunas from forested dunes. The pre-settlement sample differed in snail diversity, with 23 species compared with 9–20 species in contemporary samples. Six species were unique to the pre-settlement sample: Carychium exile, Euconulus fulvus, Gastrocopta armifera, Gastrocopta corticaria, Vallonia parvula, and Vertigo sp. The number of individual snails was much greater at the pre-settlement site when corrected for difference in size of sample area. The reason for the higher diversity, greater number of individuals, and different fauna associated with the pre-settlement area is unclear, but may be attributed to reduction and changes in the litter layer at the contemporary comparison sites caused by European earthworms. This work suggests that major erosional and depositional events occurred on Great Lakes shoreline dunes in the past and that these can be used to study post glacial mollusc faunas and past ecological processes, with some potentially significant results.
2015
McAlpine DF, Lepitzki DAW, Schueler FW, McAlpine FJT, Hebda A, Forsyth RG, Nicolai A, Maunder JE, Noseworthy RG (2015) Occurrence of the Chinese mystery snail, Cipangopaludina chinensis (Gray, 1834) (Mollusca: Viviparidae) in the Saint John River system, New Brunswick, with review of status in Atlantic Canada. BioInvasions Journal 5 (3): 149–154. doi: 10.3391/bir.2016.5.3.05
The Chinese mystery snail, Cipangopaludina [=Bellamya] chinensis, is documented for the first time in the Saint John River, New Brunswick, a watercourse which drains the largest watershed in Atlantic Canada. This is the first non-native mollusc known to be established in the Saint John River system. Although significant ecosystem effects of the species seem unlikely, possible introduction of C. chinensis via boat traffic emphasizes the need for boater education combined with monitoring of the Saint John River system for potentially more troublesome non-native species, including the zebra mussel, Dreissena polymorpha. Review of published records and museum collections for Atlantic Canada show C. chinensis has been reported from 13 freshwater wetlands and waterbodies in New Brunswick, Nova Scotia and Newfoundland since 1955. The species remains extant in at least 10 of these sites and is clearly more widespread in the region than the single previously published report would suggest.
Forsyth RG, Oldham MJ (2016) Terrestrial molluscs from the Ontario Far North. Check List 12 (3): 1881. doi: 10.15560/12.3.1881
From 2009–2014, incidental collections of terrestrial molluscs were made from the Far North of Ontario, Canada. Thirty-four species of terrestrial molluscs were identified from these collections, including one exotic species, Deroceras reticulatum, and three newly reported species from the Far North, Vallonia pulchella, Vertigo cf. genesii, and Gastrocopta similis.Vertigo cf. genesii is newly reported from Ontario. Some species have not been collected in Ontario for many years and some of these only once before. Subnational conservation ranks are discussed for higher-ranked species.
Telfer AC, 111 additional authors (2015) Biodiversity inventories in high gear: DNA barcoding facilitates a rapid biotic survey of a temperate nature reserve. Biodiversity Data Journal 3: e6313. doi: 10.3897/bdj.3.e6313
Comprehensive biotic surveys, or ‘all taxon biodiversity inventories’ (ATBI), have traditionally been limited in scale or scope due to the complications surrounding specimen sorting and species identification. To circumvent these issues, several ATBI projects have successfully integrated DNA barcoding into their identification procedures and witnessed acceleration in their surveys and subsequent increase in project scope and scale. The Biodiversity Institute of Ontario partnered with the rare Charitable Research Reserve and delegates of the 6th International Barcode of Life Conference to complete its own rapid, barcode-assisted ATBI of an established land trust in Cambridge, Ontario, Canada. The existing species inventory for the rare Charitable Research Reserve was rapidly expanded by integrating a DNA barcoding workflow with two surveying strategies—a comprehensive sampling scheme over four months, followed by a one-day bioblitz involving international taxonomic experts. The two surveys resulted in 25,287 and 3,502 specimens barcoded, respectively, as well as 127 human observations. This barcoded material, all vouchered at the Biodiversity Institute of Ontario collection, covers 14 phyla, 29 classes, 117 orders, and 531 families of animals, plants, fungi, and lichens. Overall, the ATBI documented 1,102 new species records for the nature reserve, expanding the existing long-term inventory by 49%. In addition, 2,793 distinct Barcode Index Numbers (BINs) were assigned to genus or higher level taxonomy, and represent additional species that will be added once their taxonomy is resolved. For the 3,502 specimens, the collection, sequence analysis, taxonomic assignment, data release and manuscript submission by 100+ co-authors all occurred in less than one week. This demonstrates the speed at which barcode-assisted inventories can be completed and the utility that barcoding provides in minimizing and guiding valuable taxonomic specialist time. The final product is more than a comprehensive biotic inventory—it is also a rich dataset of fine-scale occurrence and sequence data, all archived and cross-linked in the major biodiversity data repositories. This model of rapid generation and dissemination of essential biodiversity data could be followed to conduct regional assessments of biodiversity status and change, and potentially be employed for evaluating progress towards the Aichi Targets of the Strategic Plan for Biodiversity 2011–2020.
Forsyth RG, Oldham MJ, Snyder E, Schueler FW, Layberry R (2015) Forty years later: distribution of the introduced Heath Snail, Xerolenta obvia in Ontario, Canada (Mollusca: Gastropoda: Hygromiidae). Check List 11 (4): 1711. doi: 10.15560/11.4.1711
Xerolenta obvia is a Central European land snail that is introduced to Ontario, Canada, where it was first recorded in the literature in 1975 from a single population at Bethany (City of Kawartha Lakes). Over the four decades since that publication, additional records have been found clustered near Bethany, within the City of Kawartha Lakes, and in neighbouring Peterborough County and the Regional Municipality of Durham. Two distant sites are also now known, one in rural Ottawa (southeastern Ontario), and one at Windsor, Essex County (southwestern Ontario).
Forsyth RG, Lepitzki DAW (2015) Terrestrial snails (Gastropoda: Pulmonata) from Writing-on-Stone Provincial Park, Alberta, Canada. Check List 11 (3): 1636. doi: 10.15560/11.3.1636
There are few studies on the terrestrial molluscs of the Great Plains of Alberta. Nearly all previously published surveys have focused on the faunas of the Rocky Mountain Foothills, the Cypress Hills, the Parklands of Central Alberta, or the boreal forests. By means of hand-picking and litter samples, we surveyed for the first time the terrestrial snail fauna in the Writing-on-Stone Provincial Park, southern Alberta, Canada. From among the hoodoos (weathered rock formations) and along the cliffs on the north side of the Milk River, we found terrestrial snails at four of our five sites within the hot, dry valley of the river. Our study confirms our assumptions that the terrestrial snail fauna in these hostile habitats lacks diversity.
Forsyth RG, Oldham MJ, Schueler FW (2015) Patera appressa (Say, 1821), an introduced land snail in Ontario, Canada (Mollusca: Gastropoda: Polygyridae). Check List 11 (2): 1583. doi: 10.15560/11.2.1583
The geographic distribution and habitat of Patera appressa (Say, 1821) in Canada are described. This is a North American land snail native to the southern Appalachians but it has been known, since 1994, from a cluster sites in Trenton, City of Quinte West, Ontario, where it is presumably introduced. It may no longer be extant at any of the known Ontario sites.
Forsyth RG (2015) First record of Carychium minimum Müller, 1774 in New Brunswick, Canada (Mollusca: Gastropoda: Ellobioidea). Check List 11 (1): 1511. doi: 10.15560/11.1.1511
The minute land snail, Carychium minimum Müller, 1774 is reported from New Brunswick, Canada. This new record further adds additional data to support the supposition that this introduced, European species is probably more widespread over temperate North America than currently known.
2014
Forsyth RG, Oldham MJ (2014) Distribution of Strobilops aeneus Pilsbry, 1926, in Canada, with two new Ontario records (Mollusca: Gastropoda: Strobilopsidae). Check List 10 (2): 397–401. doi: 10.15560/10.2.397
The geographic distribution of Strobilops aeneus Pilsbry, 1926, a rare species in Canada, is reviewed and all known records are mapped. Two recent records, the only ones since 1941, are reported from the province of Ontario. One of these records represents a small range extension ca. 85 km north of the closest previous site. Specimens identified as S. aeneus from New Brunswick and Nova Scotia were re-examined and found to be another species.
McAlpine DF, Forsyth RG (2014) Occurrence of the Copse Snail, Arianta arbustorum (Helicidae) on Prince Edward Island: an addition to the North American range of a purported potential pest. Northeastern Naturalist 21 (1): N5–N7. doi: 10.1656/045.021.0123 · Link to PDF
We report the first occurrence of Arianta arbustorum (Copse Snail) on Prince Edward Island, expanding its well-established presence in eastern Canada. Although listed as an exotic mollusc for which prevention of introduction to the US is a priority, the evidence from Canada to date suggests a low pest potential for this species. It is unlikely that eradication efforts in North America would be effective; thus, the Copse Snail may be a poor candidate for pest-prevention efforts in the US.
Forsyth RG (2014) First record of Deroceras invadens Reise, Hutchinson, Schunack & Schlitt, 2011 (Gastropoda: Pulmonata: Agriolimacidae) from the island of Newfoundland, Canada. Check List 10 (1): 149–150. https://doi.org/10.15560/10.1.149
The introduced European slug Deroceras invadens Reise, Hutchinson, Schunack & Schlitt, 2011 is here reported from St. John’s, Newfoundland. This new record is the first from the island of Newfoundland, the Canadian province of Newfoundland and Labrador, and from Atlantic Canada. It is the first verified record for this species living outside of greenhouses in eastern Canada.
2013
COSEWIC (2013) COSEWIC Status Appraisal Summary on the Puget Oregonian Cryptomastix devia in Canada. Committee on the Status of Endangered Wildlife in Canada, Ottawa, Ontario, Canada, xv pp. [Report by R.G. Forsyth]. Link to PDF
2012
Forsyth RG, Williston P (2012) Terrestrial snails from an urban park in Vancouver, British Columbia. The Festivus 44 (7): 77–80. Biodiversity Heritage Library
[We report on land snails found in leaf-litter samples collected in Queen Elizabeth Park, Vancouver, BC, Canada. Although no native forest remains and garden areas are with ornamental trees and shrubs (decades old and mature), we found three native species of snail. We also verify recent finds in the park of Carychium tridentatum (Risso, 1826) and Discus rotundatus (Müller, 1774) and report an additional nearby site for D. rotundatus, as well as discuss a historical record.]
2010
Grimm FW, Forsyth RG, Schueler FW, Karstad A ("2009") [2010] Identifying Land Snails and Slugs in Canada: Introduced Species and Native Genera. Canadian Food Inspection Agency, Ottawa, Ontario, Canada. [April 2010].
Grimm FW, Forsyth RG, Schueler FW, Karstad A ("2009") [2010] Identification des escargots et des limaces terrestres au Canada: Espèces introduites et genres indigènes. Agence canadienne d’inspection des alimentes, Ottawa, Ontario, Canada. [April 2010].
Lepitzki DAW, Forsyth RG, Lepitzki BM (2010) The mountainsnails of Cypress Hills, Alberta, Canada. Tentacle 18: 3–5. Link to publication
2008
Forsyth RG, Oldham MJ, Schueler FW (2008) Mollusca, Gastropoda, Ellobiidae, Carychium minimum, and Ferussaciidae, Cecilioides acicula: distribution extension and first provincial records of two introduced land snails in Ontario, Canada. Check List 4 (4): 449–452. doi: 10.15560/4.4.449
Forsyth RG (2008) First record of the European land snail Trochulus striolatus in British Columbia, Canada (Pulmonata: Hygromiidae). The Festivus 40 (6): 76–78. Biodiversity Heritage Library
The European terrestrial snail Trochulus striolatus (C. Pfeiffer, 1828), found at one locality in the city of Revelstoke, is newly reported from British Columbia, Canada. This record appears to be the first time that this species has been found in western North America.
2005
Forsyth RG (2005) Terrestrial gastropods of the upper Fraser Basin of British Columbia. Living Landscapes, Royal British Columbia Museum, Victoria, British Columbia, Canada. 26 pp. Link to PDF
Information on the terrestrial gastropod fauna is compiled from new or recent field collections, museum records and literature for that part of British Columbia encompassing the basin of the Fraser River north of about 52°N. Recent fieldwork (2001) has added many new locality records for a region that has received little study and has significantly improved the region’s representation of terrestrial gastropods in the collection of the Royal British Columbia Museum. At least 28 species belonging to 18 genera are recorded from 83 localities. All but three species were previously unreported in the literature from this area of British Columbia.
Forsyth RG (2005) Terrestrial gastropods of the Peace River–northern Rockies of British Columbia. Living Landscapes, Royal British Columbia Museum, Victoria, British Columbia, Canada. 23 pp. Link to PDF
For the first time, information on the terrestrial mollusc fauna is compiled from new or recent field collections, museum records and literature for the Alberta and Liard plateaus, the northern Rocky Mountains, and much of the Cassiar and Omineca mountains of British Columbia, an area about 265,000 km2. Based on recent fieldwork (2003–2004) many new locality records are given for a poorly known region of the province. At least 23 species belonging to 13 families and 17 genera are recorded from 77 unique localities are documented in an annotated species list. Distribution of each species within the region is mapped, and locality data is tabulated in an appendix.
Reise H, Hutchinson JMC, Forsyth RG, Forsyth TJ (2005) First records of the terrestrial slug Deroceras turcicum (Simroth, 1894) in Poland. Folia Malacologica 13 (4): 177–179. doi: 10.12657/folmal.013.017
Deroceras turcicum (Simroth) is reported from six woodland sites around Wałbrzych in southwest Poland. This extension of the species’ range to Poland was expected given the number of reports from adjacent areas of the Czech Republic. We collate these reports as well as local records of Deroceras praecox Wiktor, 1966, which is found in similar habitats. We briefly discuss the difficulty of distinguishing D. turcicum from Deroceras reticulatum (O.F. Müll.), with which it may also co-occur.
2004
Forsyth RG (2004) Gastrocopta in British Columbia (Mollusca: Pulmonata: Vertiginidae). The Festivus 36 (5): 53–55. Biodiversity Heritage Library
Gastrocopta holzingeri (Sterki, 1889), previously known in British Columbia, Canada, by unpublished 40-year-old museum records, was rediscovered in southeastern B.C. in 2002, and its habitat is described. References in the literature to G. pentodon (Say, 1822) in British Columbia are thought to be in error.
Forsyth RG (2004) Land Snails of British Columbia. Royal British Columbia Museum Handbook. Royal BC Museum, Victoria, British Columbia, Canada.
2003
Forsyth RG (2003) Northern range extension for Assiminea translucens (Carpenter, 1864). The Festivus 35 (2): 17–18. Biodiversity Heritage Library
[Assiminea translucens (= A. californica) is reported from Larcom Island, Observatory Inlet, northen British Columbia (55°23.39′N, 124°53′W). This species was previously known to occur north to Vancouver Island.]
2002
Forsyth RG ("2001") [2002] New records for land snails from the mountains of northwestern British Columbia. The Canadian Field-Naturalist 115 (2): 223–228. doi: 10.5962/p.363781
Terrestrial gastropod faunas of vast areas of northern and central British Columbia are extremely poorly known. I record new distributional records of Punctum randolphii (Dall, 1895), Microphysula cookei (Pilsbry, 1922), Pristiloma arcticum (Lehnert, 1884), Vertigo columbiana (Pilsbry & Vanatta, 1900) and other land snails from the Babine and Bulkley Ranges, Nechako Plateau, and Boundary Range of northwestern British Columbia. Records of Punctum randolphii andVertigo columbiana are the first higher elevation localities in British Columbia. Pristiloma arcticum was collected from one site in the Babine Mountains and is newly recorded from the province. Microphysula cookei, previously known only from southern British Columbia and adjacent Washington, occurs north to the Babine and Hazelton Ranges, Nechako Plateau, and Boundary Range. While sharing many species in common, the mountain snail fauna differed from that of the adjacent Bulkley Valley by the inclusion of “coastal” and montane elements. Absent from the drier valley floor, Punctum randolphii, Microphysula cookei, Vertigo columbiana and Vespericola columbianus are considered coastal species whose distributions extend far inland along climatically favourable corridors.
COSEWIC (2002) COSEWIC assessment and status report on the Warty Jumping-slug, Hemphillia glandulosa, in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa. vi + 19 pp. [Status report prepared by R.G. Forsyth and K.E. Ovaska.]
COSEWIC (2002) COSEWIC assessment and status report on the Dromedary Jumping-slug, Hemphillia dromedarius, in Canada. Committee on the Status of Endangered Wildlife in Canada, Ottawa, Ontario, Canada. [Status report prepared by K.E. Ovaska and R.G. Forsyth.]
COSEWIC (2002) COSEWIC assessment and status report on the Oregon forestsnail, Allogona townsendiana, in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa, Ontario, Canada. [Status report prepared by R.G. Forsyth and K.E. Ovaska.]
COSEWIC (2002) COSEWIC assessment and status report on the Puget Oregonian snail, Cryptomastix devia, in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa, Ontario, Canada. [Status report prepared by K.E. Ovaska and R.G. Forsyth.]
2001
Forsyth RG (2001a) Re-identification of slugs from seabird nesting burrows off the west coast of Vancouver Island. The Festivus 33 (1): 9–10. Biodiversity Heritage Library
Two species of terrestrial slugs were collected from driftwood and the nesting burrows of Leach’s storm-petrels, Oceanodroma leucorhoa (Vieillot, 1818), on a small island off the west coast of Vancouver Island in the late 1960s. The species were identified and published as Ariolimax columbianus (Gould, 1851) and Deroceras reticulatum (Müller, 1774). However, based on external morphology and anatomy of preserved material and remarks on the pigmentation of the living animal, the record is re-identified as Prophysaon foliolatum (Gould, 1851). Although incorrectly determined, the original observations of P. foliolatum from storm-petrel burrows are noteworthy because of the unusual slug–bird association and habitat.
Forsyth RG (2001) A note on the distribution of Striatura pugetensis in British Columbia. The Festivus 33 (5): 57–58. Biodiversity Heritage Library
Forsyth RG (2001) First records of the European land slug Lehmannia valentiana in British Columbia, Canada. The Festivus 33 (7): 75–78. Biodiversity Heritage Library
The synanthropic slug, Lehmannia valentiana (Férussac, 1821), is recorded in British Columbia for the first time. The identification of an adult specimen from Victoria, Vancouver Island, is confirmed by dissection. Additional records, based on undissected immature specimens, photographs and sightings, are known from nearby sites on southern Vancouver Island and from an island in the Strait of Georgia.
Forsyth RG, Forsyth T (2001) A note on Mercenaria mercenaria in British Columbia. The Festivus 33 (8): 85. Biodiversity Heritage Library
[The finding of Mercenaria mercenaria in Boundary Bay, British Columbia, Canada was mentioned in two publications in 1998 and 2000. Data are provided on this specimen and photograph of the shell is provided.]
Forsyth RG, Hutchinson JMC, Reise H (2001) Aegopinella nitidula (Draparnaud, 1805) (Gastropoda: Zonitidae) in British Columbia—first confirmed North American record. American Malacological Bulletin 16 (1/2): 65–69. Biodiversity Heritage Library
The European land snail, Aegopinella nitidula (Draparnaud, 1805), is reported for the first time from British Columbia, from three sites in the city of Vancouver. These new records are the only documentation of the species in North America, except for two old records that are probably erroneous and have been ignored in recent literature. Comparisons are made between A. nitidula and similar native and introduced species. Information about its ecology in Europe is summarized.
2000
Forsyth RG (2000) The land snail Cryptomastix germana (Gastropoda: Polygyridae) in the Queen Charlotte Islands, British Columbia: a range extension north from Vancouver Island. The Canadian Field-Naturalist 114 (2): 316–317. doi: 10.5962/p.363970
The coastal land snail Cryptomastix germana, not previously known to occur north of Vancouver Island, is reported from the Queen Charlotte Islands, British Columbia.
Reise H, Hutchinson JMC, Forsyth RG, Forsyth TJ (2000) The ecology and rapid spread of the terrestrial slug Boettgerilla pallens in Europe with reference to its recent discovery in North America. The Veliger 43 (4): 313–318. Biodiversity Heritage Library
The terrestrial slug Boettgerilla pallens Simroth, 1912, is reported from two sites on Vancouver Island, British Columbia, the first records for this Palaearctic species in America. This paper describes how to recognize the species, and summarizes European studies of its ecology. It is unusually wormlike in appearance, lives mostly underground, and occurs in a very wide range of habitats. This century the species has spread remarkably far and fast across Europe from the Caucasus. This is demonstrated by a table of first occurrences in each country, and by three case studies of spread within Great Britain, Belgium, and north-west Austria. We predict that it will spread rapidly in North America, and may already occur more widely, but there is no evidence that it will become an important pest.
1999
Forsyth RG (1999) Lindeman Lake, British Columbia, type locality of Zonitoides randolphi Pilsbry. The Veliger 42 (3): 286. Biodiversity Heritage Library
Forsyth RG (1999) Terrestrial gastropods of the Columbia Basin, British Columbia. Living Landscapes, Royal British Columbia Museum, Victoria, British Columbia, Canada. Archived here
Forsyth RG (1999) Distributions of nine new or little-known exotic land snails in British Columbia. The Canadian Field-Naturalist 113 (4): 559–568. doi: 10.5962/p.358656
Introduced species of terrestrial molluscs of British Columbia were collected between 1989 and 1998. New locality records and expanded distributions are documented for seven introduced land snails: Lauria cylindracea (Chrysalis Snail), Vallonia pulchella (Lovely Vallonia), Oxychilus alliarius (Garlic Glass-snail), O. cellarius (Cellar Glass-snail), O. draparnaudi (Dark-bodied Glass-snail), Cepaea nemoralis (Grovesnail) and Helix aspersa (Brown Gardensnail). Two other species, Vallonia excentrica (Iroquois Vallonia), and Vitrea contracta (Contracted Glass-snail) are reported from British Columbia for the first time. About 25% of the terrestrial mollusc species in the province are exotic. Mechanisms of introduction include transport on nursery plants and on salvaged bricks. Potential impacts on native fauna include predation by Oxychilus.